TYPE Costa Rica. Coto Brus, Sabalito, San Miguel, carretera entre Union y Mellizas, 2.5 km después de Unión, orillas de la Quebrada Sereno, 8°51 ' 17.0"N, 82°52'35.2"W, 994 m, bosque muy humedo premontano transicion a pluvial "supra truncos ad ripas silvarum prope San Miguel juxta flumen ad Quebrada Sereno," April 18, 2011, A. P. Karremans 3963, D. Bogarin & D. Jiménez (holotype, JBL-spirit; illustration of the plant that served as the holotype).